【作者单位】1 China–UK HUST–RRes Genetic Engineering and Genomics Joint Laboratory, Huazhong University of Science and Technology, Wuhan, Hubei, 430074, China; 2 School of Life Sciences and Center for Evolutionary Biology, Fudan University, Shanghai, 200433, China; 3 Institute of Biodiversity Science and Geobiology, Tibet University, Lhasa, 850000, China; 4 ‖Millennium Seed Bank Project, Royal Botanic Gardens, Kew, Wakehurst Place, Ardingly, West Sussex RH17 6TN, UK