【作者单位】Affiliations 1 Academy of Plateau Science and Sustainability, School of Life Sciences, Qinghai Normal University, Xining, Qinghai, 810000, China; College of Life Sciences, Beijing Normal University, Beijing, 100875, China. Electronic address: ydwang@bnu.edu.cn. 2 Academy of Plateau Science and Sustainability, School of Life Sciences, Qinghai Normal University, Xining, Qinghai, 810000, China. 3 Academy of Plateau Science and Sustainability, School of Life Sciences, Qinghai Normal University, Xining, Qinghai, 810000, China; College of Life Sciences, Beijing Normal University, Beijing, 100875, China. 4 Northwest Institute of Plateau Biology, Chinese Academy of Sciences, Xining, Qinghai, 810008, China. 5 Department of Life Sciences, College of Science, Tibet University, Tibet University, Lhasa, Tibet, 850000, China. 6 Academy of Plateau Science and Sustainability, School of Life Sciences, Qinghai Normal University, Xining, Qinghai, 810000, China; MOE Key Laboratory of Cell Activities and Stress Adaptations, School of Life Sciences, Lanzhou University, Lanzhou, 730000, China. Electronic address: qiuqsh@lzu.edu.cn.