【作者单位】School of Ecology and Environment，Tibet University，Lhasa 850000，China；CAS Key Laboratory of Aquatic Botany and Watershed Ecology，Wuhan Botanical Garden，Chinese Academy of Sciences，Wuhan 430074，China；University of Chinese Academy of Sciences，Beijing 100049，China；CAS Key Laboratory of Aquatic Botany and Watershed Ecology，Wuhan Botanical Garden，Chinese Academy of Sciences，Wuhan 430074，China；University of Chinese Academy of Sciences，Beijing 100049，China；Department of Biosciences，Rice University，Houston，TX 77005，USA；Key Laboratory of Alpine Vegetation Ecological Security，Institute of Tibet Plateau Ecology，Tibet Agriculture and Animal Husbandry University，Nyingchi 860000，China；Resource and Environment College，Tibet Agriculture and Animal Husbandry University，Nyingchi 86000，China；School of Ecology and Environmental Science，Yunnan University，Kunming 650504，China；CAS Key Laboratory of Aquatic Botany and Watershed Ecology，Wuhan Botanical Garden，Chinese Academy of Sciences，Wuhan 430074，China；Department of Biosciences，Rice University，Houston，TX 77005，USA
