【作者单位】a Key Laboratory of Eco-environments in Three Gorges (Ministry of Education), Chongqing Engineering Research Centre for Sweetpotato, School of Life Sciences, Southwest University, Chongqing 400715, China b College of Pharmaceutical Sciences, Southwest University, Chongqing 400715, China c Agricultural and Animal Husbandry College, Tibet University, Nyingchi of Tibet 860000, China